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This article describes the sound changes from Proto-Nautli to Ancient Zana. The stages are named for the most distinguishable sound changes taking place in that time.

Loss of Pharngealization Contrast Edit

V → Vˤ / {Cˤ, ʕ, ʕʷ}_
Cˤ → C
ɮ → z
ʕ → ∅
h → ∅
ʕʷ → w
{i ɪ e ɛ u ʊ o ɔ a ɐ}ˤ → {ɘ ɜ a ɐ ɘ ɜ a ɐ a ɐ}

Loss of Retroflexes Edit

{ʈ ʂ} → {t s}
ɻ → l

Loss of Labialization Contrast Edit

{i ɪ e ɛ ɘ ɜ a ɐ} → {y ʏ ø œ ɵ ɞ ɒ ɐ̹} / {Cʷ, w}_
Cʷ → C
w → ∅

Simplification of Vocalic System Edit

{ɘ ɜ} → {e ɛ}
{y ʏ ø œ ɵ ɞ} → {u ʊ o ɔ o ɔ}
{ɒ ɐ̹} → {o ɔ}